Dharragarra

Dharragarra Temporal range: Late Cretaceous (Cenomanian), 100.2–96.6 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Monotremata
Family:	Ornithorhynchidae (?)
Genus:	†Dharragarra Flannery, 2024
Species:	†D. aurora
Binomial name
†Dharragarra aurora Flannery et al., 2024

Dharragarra (meaning "platypus" in the Gamilaraay language) is an extinct genus of monotreme mammal from the Late Cretaceous (Cenomanian) Griman Creek Formation of Australia. The genus contains a single species, D. aurora, known from a partial left mandibular ramus. Dharragarra was likely more closely related to the living platypus than many other monotremes of the Cretaceous.

Discovery and naming

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Dharragarra type locality near Lightning Ridge, Australia

The Dharragarra holotype specimen, AM F97262, was discovered in 2002 in sediments of the Griman Creek Formation (Wallangulla Sandstone Member) near Lightning Ridge, New South Wales, Australia. The specimen consists of a partial left horizontal mandibular ramus.^[1] This bone was first mentioned by Anne Musser in part of a 2013 publication, where it was identified as a steropodontid.^[2] In a 2022 monotreme evolution review, Flannery et al. alluded to it as an unnamed new genus of stem-ornithorhynchid.^[3]

In 2024, Flannery et al. described Dharragarra aurora as a new genus and species of early monotreme based on these fossil remains. The generic name, Dharragarra, is a word that means "platypus" in the Gamilaraay, Yuwaalaraay and Yuwaalayaay languages, referencing this taxon's resemblance to the extant lineage. The specific name, aurora, is a Latin word meaning "dawn", considering the taxon's position at the beginning of monotreme evolution.^[1]

Several other monotremes are known from the Griman Creek Formation, including Opalios and Parvopalus—which were described in the same publication as Dharragarra—as well as Kollikodon, Steropodon, and Stirtodon.^[1][4]

Classification

Dharragarra is one of the oldest members of the platypus and echidna lineage, being closer to them than the coeval ornithorhynchoid Opalios. Due to the close resemblance of its jaw to that of a platypus, Dharragarra is considered one of the oldest members of the platypus stem lineage, although it likely predates the divergence of platypuses and echidnas.^[1]

References

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Mammalia

Kingdom: Animalia Phylum: Chordata Clade: Synapsida Clade: Therapsida Clade: Cynodontia Clade: Mammaliaformes
Synapsida see Synapsida Cynodontia see Cynodontia Mammalia see below↓

Mammalia
Australosphenida? †Ambondro †Tendagurutherium? Monotremata? †Henosferidae Asfaltomylos Henosferus †Ausktribosphenidae Ausktribosphenos Bishops? Kryoparvus? Monotremata †Kollikodontidae? Kollikodon Kryoryctes? Sundrius †Steropodontidae Parvopalus? Steropodon †Teinolophidae Stirtodon? Teinolophos Ornithorhynchoidea †Opalios Ornithorhynchidae †Dharragarra? †Monotrematum? †Obdurodon Ornithorhynchus †Patagorhynchus? Tachyglossidae †Megalibgwilia †Murrayglossus Tachyglossus Zaglossus †Allotheria? see Allotheria includes multituberculates Theriimorpha see below↓

Theriimorpha
Theriimorpha †Acinacodus †Amphidon †Amphilestes †Aploconodon †Chaoyangodens †Comodon †Condorodon †Dyskritodon? †Fruitafossor? †Hakusanodon †Indotriconodon †Juchilestes †Jueconodon †Kemchugia †Kryptotherium †Liaoconodon †Liaotherium †Manchurodon †Nakunodon †Phascolotherium †Sangarotherium †Tendagurodon †Allotheria? Monotremata? †Gobiconodontidae Fuxinoconodon Gobiconodon Hangjinia Huasteconodon? Meemannodon Repenomamus Spinolestes †Jeholodentidae Jeholodens Yanoconodon †Klameliidae? Ferganodon Klamelia †Triconodontidae Alticonodon Arundelconodon Astroconodon Corviconodon Eotriconodon Jugulator Meiconodon Priacodon Triconodon Trioracodon Victoriaconodon Volaticotheria? †Volaticotheria Argentoconodon Ichthyoconodon Triconolestes? Volaticotherium †Tinodontidae Gobiotheriodon? Tinodon Trishulotherium Yermakia Trechnotheria Monotremata? †Zhangheotheriidae Anebodon Kiyatherium Maotherium Ningchengodon Origolestes Zhangheotherium †Spalacotheriidae Akidolestes Aliaga Cifellitherium Heishanlestes Infernolestes Lactodens Shalbaatar Spalacolestes Spalacotheridium Spalacotherium Spalacotheroides Symmetrodontoides Symmetrolestes Yaverlestes Cladotheria see below↓

Cladotheria
†Dryolestida Brancatherulum? Dorsetodon Drescheratherium Euthlastus Henkelotherium Paurodon Tathiodon Dryolestidae Achyrodon Amblotherium Anthracolestes Beckumia Crusafontia Dryolestes Groebertherium? Guimarotodus Hercynodon Krebsotherium Lakotalestes? Laolestes Minutolestes Phascolestes Portopinheirodon Treslagosodon †Meridiolestida Amarillodon Austrotriconodon Bondesius Casamiquelia Cronopio Groebertherium? Lakotalestes? Leonardus Necrolestes Paraungulatum Quirogatherium Solanutherium Mesungulatoidea Coloniatherium Mesungulatum Orretherium Peligrotherium Reigitherium †Donodontidae Amazighodon Anoualestes Donodon Stylodens Prototribosphenida †Abelodon †Amphibetulimus †Amphitherium †Arguimus †Brancatherulum? †Chunnelodon †Magnimus †Minimus †Mozomus †Nanolestes †Palaeoxonodon †Vincelestes Monotremata? †Peramuridae Afriquiamus? Kouriogenys Peramuroides Peramus Tendagurutherium? Tribosphenida / Boreosphenida †Aegialodon †Argaliatherium †Brasilestes †Carinalestes †Eomaia †Holoclemensia †Hypomylos †Juramaia †Kermackia †Kielantherium †Pappotherium †Picopsis †Tirotherium †Tribactonodon Theria †Patagomaia Eutheria see Eutheria includes placentals Metatheria see Metatheria includes marsupials

Other taxa
Incertae sedis †Atlasodon †Chronoperates †Microderson †Thereuodon †Tingamarra Saint Bathans Mammal UA 8699 Other taxa Prototheria Yinotheria Eutriconodonta Amphilestidae Theriiformes Pantotheria Symmetrodonta Eupantotheria Paurodontidae Amphitheriidae Marsupionta